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  1. Identify the habitat characteristics useful for predicting the occurrence of harvestmen species in caves and the epigean environment.
  2. Evaluate the intensity of fluctuating assymetry as an indicator of habitat quality in the cave habitats as compared to epigean habitats with different intensity of human impact. Differences in fluctuating assymetry at different sites could be interpreted as an indication of responses to habitat suitability.

These could yield important information about factors that influence: i) species occurrence in epigean and cave habitats and ii) factors involved in caves colonization by harvestmen.